Thanks again to all. It has been an education for me. In recording the reed frequency deviation from A=440 I adapted another spreadsheet normally used for equal temperament accordion work where G# and Ab, and D# and Eb, are the same. This initial table of recorded deviations ( attached ) shows well the differences that mean tone produces. It is not however clear to me what the fundamental tuning frequency would have been.
Matthew's Wheatstone as found.pdf